THE JOURNAL OF EXPERIMENTAL MEDICINE

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THE JOURNAL OF EXPERIMENTAL MEDICINE Senior Editors Marlowe S.Tessmer Heather L. Van Epps Scientific Editors Teodoro Pulvirenti Jennifer Smith-Garvin Editors Jean-Laurent Casanova Adolfo Garcia-Sastre David Holtzman Lewis L. Lanier William A. Muller Carl Nathan Michel Nussenzweig Anne O’Garra Alexander Rudensky Alan Sher Sasha Tarakhovsky Andreas Trumpp David Tuveson Editor Emeritus Alan N. Houghton Manuscript Coordinators Meagan Brooks and Sylvia F. Cuadrado phone (212) 327-8575 fax (212) 327-8511 email: [email protected]

Preflight Editor Rochelle Ritacco Copy Editors Brianna Caszatt and Maya Frank-Levine Production Editor Shauna O’Garro Production Coordinator Camille Clowery Electronic Production Coordinator Erinn A. Grady

Advisory Editors Shizuo Akira Kari Alitalo Frederick W. Alt K. Frank Austen Albert Bendelac Michael J. Bevan Christine A. Biron Christian Bogdan Hal E. Broxmeyer Meinrad Busslinger Arturo Casadevall Ajay Chawla Yongwon Choi Robert L. Coffman Daniel J. Cua Myron I. Cybulsky Riccardo Dalla Favera Glenn Dranoff Michael Dustin Douglas T. Fearon Vincent A. Fischetti Richard A. Flavell Patricia Gearhart Ronald N. Germain Christopher Goodnow Siamon Gordon Or Gozani Sergio Grinstein Philippe Gros Kristian Helin Kristin A. Hogquist Chyi Hsieh Christopher A. Hunter Kayo Inaba Gerard Karsenty Jay Kolls Paul Kubes Vijay K. Kuchroo

Ralf Kuppers Tomohiro Kurosaki Bart N. Lambrecht Klaus F. Ley Yong-Jun Liu Clare Lloyd Tak Mak Bernard Malissen James S. Malter Philippa Marrack Diane Mathis Ira Mellman Matthias Merkenschlager Sean J. Morrison Muriel Moser Christian Münz Cornelis Murre Benjamin G. Neel Michael Neuberger Victor Nussenzweig John J. O’Shea Paul H. Patterson Fiona Powrie Lluis Quintana-Murci Klaus Rajewsky Gwendalyn J. Randolph Anjana Rao Jeffrey Ravetch Sergio Romagnani Nikolaus Romani David L. Sacks Shimon Sakaguchi Federica Sallusto Matthew D. Scharff Olaf Schneewind Stephen P. Schoenberger Hans Schreiber Gerold Schuler Robert A. Seder

Rafick-P. Sékaly Charles N. Serhan Nilabh Shastri Ethan M. Shevach Roy L. Silverstein Jonathan Sprent Louis M. Staudt Janet Stavnezer Andreas Strasser Toshio Suda Stuart Tangye Steven L.Teitelbaum Thomas J.Templeton Kevin J.Tracey Giorgio Trinchieri Shannon Turley Marcel R.M. van den Brink Ulrich von Andrian Harald von Boehmer Christopher M.Walker Raymond M.Welsh E. John Wherry Linda S.Wicker Ian Wilson Thomas Wynn

Monitoring Editors Marco Colonna Jason Cyster Stephen Hedrick Andrew McMichael Luigi Notarangelo

Consulting Biostatistics Editors Glenn Heller Madhu Mazumdar

Copyright to articles published in this journal is held by the authors. Articles are published by The Rockefeller University Press under license from the authors. Conditions for reuse of the articles by third parties are listed at http://www.rupress.org/terms Print ISSN 0022-1007 Online ISSN 1540-9538

EDITOR-IN-CHIEF

Linda J. Miller FINANCE DIRECTOR

Raymond T. Fastiggi BUSINESS DEVELOPMENT DIRECTOR

Gregory Flynn Malar ELECTRONIC PUBLISHING AND PRODUCTION DIRECTOR

Robert J. O'Donnell ADVERTISING SALES DIRECTOR

Lorna Petersen COMMUNICATIONS MANAGER

Rita Sullivan King OFFICE ADMINISTRATOR

JoAnn Greene ASSISTANTS TO THE FINANCE DIRECTOR

Sarah S. Kraft Ann Travers BUSINESS DEVELOPMENT SPECIALIST

Colleen M. Smith MARKETING ASSOCIATE

Laraine Karl

The Journal of Experimental Medicine (ISSN 0022-1007) is published every four weeks (1 volume, 13 issues) by The Rockefeller University Press, 1114 First Avenue, New York, NY 10065-8325. Periodical postage paid at New York, NY and additional mailing offices. 2014 Subscription Rates: Institutional Rates: Tier 1 Online $1,455 Print+Online $3,300

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